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The American Kestrel and Iowa’s Nest Box Program

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Species Description: American Kestrel Falco sparverius

 

There is a lot to learn about the American kestrel, including the fact that it is the smallest and most colorful member of the falcon (Falconidae) family in North America, and with the exception of the Seychelles kestrel, the world. male kestrel image

Click on any of the links below to learn more. 

Species

     Ecological importance and value

     Identification Tips

     Characteristics and Behaviors

Geographic range

     Habitat

     Winter movement and migration

Diet/food habits

Pair and reproduction

     Site selection

     Eggs

     Incubation 

     Nesting period

     Post-fledgling period

Life Expectancy

Species

The American kestrel (Falco sparverius) is the smallest and most colorful member of the falcon (Falconidae) family in North America, and with the exception of the Seychelles kestrel, the world.  It is roughly the size of a robin or blue jay.

The American kestrel was once called the “sparrow hawk.” However, the name was changed because the diet of a Falco sparverius is not even close to being exclusively made up of sparrows.  And, technically, the kestrel is a falcon not a hawk.  Falcons have long, pointed wings, rapid wing strokes, and they usually do not soar as much as hawks.

The name kestrel is likely derived from the Old French word “cresserelle,” which means crest and refers to the reddish-brown back and crown patch.

Because of their choice of food, kestrels are abundant in agricultural areas characterized by scattered woodlots and trees, shelterbelts, meadows, highway rights-of-way, pastures, and hay fields.  Kestrels are frequently seen sitting on power lines along highways or hovering above the grassy roadside ditches.  From their location they dive to the ground to catch their prey.

Ecological Importance and Value

The American kestrel is very important ecologically in controlling creatures humans usually consider a nuisance and destructive (rodents and large insects).

As attractive members of the falcon family, they are an interesting and aesthetically pleasing component of our natural world, especially to those who take the time to enjoy their hovering and diving techniques.

Identification tips

The average kestrel weighs anywhere from 3-6 ounces (females are slightly larger than males), is 9-12 inches long and has a wingspan of 20-24 inches.  Kestrels have been clocked at speeds of nearly 40 mph.

Kestrels have long, narrow, pointed wings; a short, dark, hooked and notched beak; dark eyes; and unfeathered legs.   

Head shot of kestrel

Photo credit:  Used with permission of the Chipper Woods Bird Observatory

The American kestrel is easily identified by two distinct black streaks (moustaches or sideburns) on each side of the head that contrast with the white throat and cheeks. It is the only North American raptor with circular nostrils. 

The eyes of the kestrel face forward, providing the bird great depth perception and enabling it to prey on its favorite foods.  The kestrel’s eyes are far superior to the human eye.  They are sensitive to light in the near ultra-violet spectrum and can see details at greater distances, allowing the bird to see the trail of a rodent on the grass. 

The voice is a shrill, high pitched, “klee” call in three to six short, rapid bursts.

male kestrel perched on fence post

Photo credit:  Courtesy of the Cornell Lab of Ornithology

Sexes are distinguished by their coloration.

Adult males

The adult males have a rust patch on their crown; rust nape, breast, and back; rust tail with a broad black band midway down and a narrow white band at the end; pale belly; blue-gray wing coverts; dark flight feathers with pale spots midway on the feathers creating a “string of pearls”; and black spots on scapulars, wing coverts and flanks.

  adult female kestrel photo

Photo credit: Courtesy of the USGS Patuxent Wildlife Research Center

 Adult females

The adult female kestrel can be identified by the pale buff breast streaked with brown; rust-born nape, back and wing coverts; back and wing coverts barred heavily with black; and rust-brown tail with numerous dark bars of even width and a narrow white band at the end.

juvenile kestrel

Photo credit: Courtesy of the University of Minnesota’s Raptor Center

 Juvenile birds

Both sexes retain juvenile wing and tail feathers until their second summer, molting them after their first breeding season.  Immature males up to a year old can be recognized by the rust-brown back barred heavily with black and a streaked breast, and females by a more poorly defined band midway on the tail. 

Characteristics/Behaviors

The kestrel has two unique characteristics that set it apart from other diurnal (daytime active) raptors or birds of prey, such as hawks, eagles and falcons. 

First, the kestrel nests primarily in tree cavities that have been excavated by woodpeckers.  This differs from other diurnal raptors that build open platform nests.

Second, a kestrel hunts from a conspicuous perch or by hovering in the air, then diving down to catch its prey on the ground. The flight is buoyant, graceful and rapid, like a large swallow. Watching kestrels hunt can be an interesting experience. 

The American kestrel is, for the most part, not a social bird.  During the mating season, males and females pair up and have joint territories.  Presumably, the pair or the male defends the territory.  The function of the territory may not be so much to ensure mating as to maintain a pair bond during the nesting season when the male is needed to help rear the offspring.

Geographic Range

The kestrel is widely distributed throughout the United States.  American kestrels live in North, Central and South America from the tree line boundary in Alaska and Canada south to Tierra del Fuego, Argentina.  The bird can also be found in the West Indies, the Juan Fernandez Islands and Chile. 

In 1982 the U.S. Fish and Wildlife Service researchers estimated the population of the American kestrel was 2.4 million birds, and appeared to be stable or even increasing throughout its range.  Little recent information has been available to assess the current population status throughout most of its range. 

Habitat

The American kestrel nests in tree cavities, woodpecker holes, crevices of buildings, holes in banks, and nest boxes.  The kestrel is highly adaptable and lives just about everywhere as long as there is some open ground for hunting and places on which to perch (e.g. telephone or fence wires).

While they prefer open country, kestrels will inhabit unforested mountainsides up to 1300 feet, grasslands, savannas, deserts, farmlands, some coastal regions, and even suburban and urban environments.

The presence of trees (alive or dead) that contain adequate nesting cavities is an important component of the kestrel’s habitat.  The other two basic components of a suitable habitat are: (a) an adequate amount of open pasture-like area of short vegetation with scattered perch sites; and (b) an adequate prey supply.

The territory size of an American kestrel varies dependent upon habitat quality (e.g., prey abundance and nest site availability). Researchers have reported territory size ranging from 124 to 783 acres, with an average of one breeding pair per mile.

Winter Movement and Migration

In winter, males and females separate territories, and the sexes have different habitat preferences.  Males defend habitat with dense vegetation, whereas females select more open areas.

Those birds in the northern most parts of the geographical range migrate, while other populations are less migratory. Populations south of approximately 35°N (same latitude as Memphis,Tenn.) are, for the most part, permanent residents.

While information on migratory movement in Iowa is not well known, band returns from Iowa-banded kestrels provide some insight.  It appears that Iowa's kestrel population probably contains a mixture of birds that nest in Iowa but winter to the south, birds that winter in Iowa but nest farther north, and those that are year-round residents.

female kestrel sitting on fence post

Diet/Food Habits

American kestrels are classified as generalist predators.  Its diet varies, but sparrows and other small nesting birds make up a fair percentage (10 percent) of its prey.  Kestrels have been observed feasting on Red-winged blackbirds and Common Grackle nestlings, which they take directly from the nest after having observed the dutiful parents carrying food to the nest for their young. The starling also ranks high on the kestrel’s menu. 

In addition to eating small birds, kestrels prey on small mammals such as bats, shrews, rats, gophers, young ground squirrels, meadow voles, deer mice, and young cottontail rabbits.  About 70 percent of the kestrel’s diet consists of mammals.

Kestrels also consume large insects or invertebrates (20 percent of diet); grasshoppers, crickets, beetles and cicadas are most favored.  Other known prey items include worms, dragonflies, caterpillars, flies, and spiders.

The remaining 1 percent of the kestrel’s diet consists of reptiles and amphibians, such as snakes, sandpiper chicks, lizards, and scorpions.

In the summer, American kestrels hunt in the early morning and evening eating large insects.  Kestrels apparently are able to capture insects more readily then other prey items. During winter, they hunt throughout the daylight hours and eat small mammals. 

Kestrels obtain an adequate supply of water from the prey items consumed and therefore do not require a free source of water.

Kestrels use a variety of prey capture techniques including perch hunting, hover hunting, and foraging on foot.  Prey are typically captured with the feet and killed with the beak, although small insects are often captured and carried with the beak.

Pairing and Reproduction

American kestrels are typically monogamous (mate with one bird at a time). Pair bonding is strong and often is permanent.

Pairing begins up to six weeks before egg laying. Pair formation is initiated by the male establishing a nesting territory and female associating and hunting with the territorial male.  Males often select the same territory that was used the previous year. The female may move among several territorial males before choosing a mate.

The male, or sometimes the female, will try to attract or court a potential mate's attention by exhibiting a series of power dives from high above the territory.  During the dive-display, the male performs a series of climbs and dives with three to five high-pitched notes uttered near the highest point of each ascent. The dive consists of a vertical drop of 33 to 66 feet, and frequently occurs directly above the perched female. 

Pairs form, courtship feeding (when the male presents food to the female) occurs and the pair mate frequently until egg laying.

Nest Site Selection

Both sexes have been observed searching for suitable nest sites, however, it is primarily initiated by the male. American kestrels are almost exclusively cavity nesters and will use a natural hole in a tree, a woodpecker's hole, a nest box, a cavity in a bank or cliff, or an enclosed space in a building. On rare occasions, kestrels may use an old stick nest of another bird, especially the enclosed nests of magpies. They may even evict current residents of desirable nest sites.

kestrel egg

American kestrel egg

Photo credit: Courtesy of the USGS Patuxent Wildlife Research Center

Eggs

Eggs are deposited directly on any debris present on the cavity floor; no nesting material is brought into the cavity.  Egg color ranges from white to cream, or pale pink to reddish-brown, usually covered with reddish-brown or gray spots/specks.

Generally, four to five eggs are laid at one- to two-day intervals.  Laying dates vary by geographical area of the world.  In Iowa, egg laying normally begins in late April to early May.  If the first nesting is unsuccessful, kestrels often renest.

  Femal kestrel incubating eggs

American kestrel incubating eggs in a nest box.

Photo credit: Courtesy USGS, Northern Prairie Wildlife Resource Center

Incubation

Incubation generally begins with the second to the last egg laid, and lasts 29 to 31 days. The female does the majority of the incubation, with the male providing her with food. Males have been known to occasionally assist with incubation by sitting on the eggs. Both sexes have brooding patches.

Nestling Period

Hatching occurs over a period of three to four days.  The young nestlings are pink skinned and covered with short, silvery down at first, but become well feathered by 20 days of age.

Once hatched, kestrel chicks are tended by both parents. The female broods and feeds the young for about the first 10 days, and the male is the primary hunter. 

The female leaves the nest one to two weeks after the eggs hatch and assists the male with feeding the young.  The young grow rapidly, attaining their adult weight in 16 to 17 days.  Fledging (leaving the nest) occurs at 28 to 31 days after hatching, but the young will return to the nest at night to roost. 

Post-fledgling Period

After nest departure, the siblings and parents remain as a unit for some time, often hunting together.  The young remain dependent on their parents for food for two to three weeks after leaving the nest. The survival rate of chicks is about 50 percent under natural conditions, but is usually higher under better conditions (e.g., human-provided nesting boxes.)

Life Expectancy

The average life expectancy of an American kestrel is approximately 15 months.  They can live up to six years in the wild, but most suffer a fatality in the second year of life, usually in the fall.  In captivity, kestrels have lived from 14 to 17 years.


Now that you've learned about the uniqueness of the American kestrel, why not get involved in the Iowa's American kestrel nest box program.  To learn more click, on the program link. 

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